Pictured: A flash is seen from an explosion that struck the Engels-2 airbase near the Russian city of Saratov in December, leaving two long-range bombers damaged
Ukrainian commandos are also believed to have been involved in the recapturing of Snake Island, a Ukrainian island located in the Black Sea.
Snake Island was captured by Russian forces on the first day of Putin's invasion, when one border guard in its garrison gained near-mythic status.
Moments before two Russian warships attacked the island, the crews hailed the garrison telling them to surrender.
The border guard responded: 'Russian warship, go f**k yourself.' His remark went viral online and became a rallying cry across all of Ukraine - and even prompted the Ukrainian government to create a postage stamp honouring the soldiers there.
From that point on, the island gained a symbolic significance, and not just for its vital tactical position, and on July 1, Ukraine was able to push Russian forces off the landmass once they were in-range of US-provided HIMARS missile systems.
However, as Russia retreated, two Su-30 fighter jets bombed the island with phosphorus bombs, perhaps as a scorched-earth tactic to destroy any weapons and equipment left behind to prevent them from falling into Ukrainian hands.
This led to Ukrainian military officials suspecting that Russia may have mined the island.
Therefore, naval special forces units were sent in.
Divers from Ukraine's SOF are understood to have approached the island underwater, checking for mines, before other forces moved in.
In this photo provided by the Ukrainian Defence Ministry Press Office on Thursday, July 7, 2022, Ukrainian soldiers install the state flag on Snake island after recapturing it from Russia
Since the war began, there have been a suspicious number of assassinations of high-profile Russian officials, either in Russia itself or in occupied territory.
One of the objectives of Ukraine's special forces since the war broke out has been to organise resistance movements in towns and cities captured by Russia, and these movements are suspected of being behind such attacks.
A number of car bombs have been reported, in which officials - such as Russian-installed mayors - have been blown up.
One high-profile attack was the assassination of Darya Dugina in Bolshie Vyazyomy, on the outskirts of Moscow, on August 20, 2022.
Dugina was the daughter of Aleksandr Dugin, a supporter of Vladimir Putin and a far-right political philosopher, who has been a key supporter of the invasion of Ukraine.
She is understood to have shared her father's views.
Dugina's car exploded at around 9.45pm. Reports suggested that her father had been due to travel with her, but switched cars at the last minute. Investigators later said they found an explosive device fixed to the underside of her car.
Two days after the attack, the Russian Federal Security Service (FSB) claimed the Ukrainian special forces were behind the killing.
Pictured: Darya Dugina, 29, was killed in a car bomb attack in Moscow in August 2022
Pictured left: Flames are seen from the car that was blown up in August 2022, killing Darya Dugina.
Right: The burnt-out wreckage of her car is seen after the attack
Russian SOF
While Ukraine's special forces have set about covertly wreaking havoc behind enemy lines, and on Russia's early attempts to seize Kyiv, the role of Russia's Spetsnaz have - from what we know - been far less subtle or effective.
According to American documents that were leaked online last month, Russia's senior commanders ordered their elite forces into direct combat in the early phases of the war- not trusting their conventional fighters to be effective on their own.
As a, showed before-and-after satellite imagery of a base used by the 22nd Separate Spetsnaz Brigade.
The images of the base in southern Russia reveal that 'all but one of five Russian Separate Spetsnaz Brigades that returned from combat operations in Ukraine in late summer 2022 suffered significant losses,' The Post reports.
The satellite images show the base in November 2021, and again a year later. The earlier image shows a well populated base full of vehicles, while the second is far more empty - which the US intel concluded shows a state of extreme depletion.
The assessment says the 22nd Separate Spetsnaz Brigade as well as two other units have suffered an estimated 90 to 95 per cent attrition rate, The Post says.
Russian spetsnaz officers march during the general rehearsals of the Victory Day Parade in front of the Kremlin at Red Square, on May 7, 2021
It takes four years for Russia to train a Spetsnaz soldier, according to the documents seen by the American publication, which concludes that it could take as long as a decade for Moscow to replenish its ranks of these elite units.
Observers have expressed their shock over the tactic.
Rob Lee, a Russia military expert and senior fellow with the Foreign Policy Research Institute, told The Post that by pushing special units to the front lines, Russian commanders have burned through them at an alarming rate.
According to Sibylline's Alexander Lord, while the structure of Ukraine's special forces is now similar to those in the west, Russia takes a different approach.
'Russia's Spetsnaz 'special designation' forces have a particular reputation, which often leads to direct comparisons with the UK's SAS or US Delta Force special forces,' he explains to MailOnline.
'However, the organisation of the Spetsnaz is not directly comparable to the Western concept of special forces.'
One such difference, he says, is the fact that Russian conscripts form a 'significant contingent' of Russia's Spetsnaz units.
'The majority of Spetsnaz units are essentially good light infantry and reconnaissance forces, rather than top-tier special forces akin to the SAS,' Lord says.
Therefore, Spetsnaz is not one single elite force, but instead should be viewed as 'highly diverse units that form part of a complex network of competing elite special forces, largely borne of the Kremlin's desire to avoid over reliance on a single force.'
On account of different factions forming between Russia's supposed elite units, they often compete with each other for resources, or even with other Russian units such as the FSB's commando forces or the Foreign Intelligence Service's Zaslon unit.
Lord says that Russia has deployed many of its elite forces, including Spetsnaz and VDV - the Russian Airborne Forces (which also suffered huge casualties in the early days of the war) - to prop up its front-line infantry.
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'Moscow also did this in Chechnya and Georgia, and in many instances it has led to highly wasteful deployments of some of Russia's most competent forces,' he says.
Pointing to Bakhmut as an example, he said the defensive efforts there by poorly trained infantry are becoming increasingly reliant on elite VDV units.
General Colonel Mikhail Teplinsky, commander of the VDV, was recently appointed deputy commander of Russian operations in Ukraine - illustrating the growing importance of the
Lord says the distinction 'between Russia's elite forces are becoming increasingly meaningless' as Putin's generals become more reliant on poorly trained reservists.
These, he says, are being used 'to plug personnel gaps across the full-breadth of the Russian military - including its special forces.'
One such example is the well respected 155th Naval Infantry Brigade, which was almost entirely wiped out in Ukraine and reconstituted with poorly trained troops.
Lord says: 'This highly attritional war has seen a large proportion of Russia's well-trained units steadily wiped out and degraded, and the same has been true of its special forces.'
Meanwhile, the activities of Russia's 'true' special forces units that are more akin to western units and those formed by Ukraine remain clouded in secrecy.
Wagner PMC
One group that has come out of the shadows since the invasion was launched is the notorious Wagner Private Military Company.
The war has seen the mercenary group - seen as the de fato army of President Putin - shift from a secretive Moscow unit allegedly hired to carry out Putin's dubious activities in Syria and Africa (where Russia has military and mining interests) to a mainstream militia widely talked about in the media.
And while Wagner had previously been seen as an off-the-books special forces unit doing the clandestine bidding of Putin around the world, they too have been used to prop-up ordinary Russian units as they take heavy casualties in Ukraine.
The group was first known to be active in Ukraine in 2014 during Russia's annexation of the Crimean peninsula, where they operated with Moscow's own units.
Visitors wearing military camouflage stand at the entrance of the 'PMC Wagner Centre', which is associated with businessman and founder of the Wagner private military group Yevgeny Prigozhin, during the official opening of the office block during National Unity Day, in St.
Petersburg, Russia, on November 4, 2022
They were soon deployed to the Donbas, where a conflict was raging between the Ukrainian government and pro-Russian forces. There, they were able to help the Russian sympathisers destabilise local governments and take control of towns.
The PMCs conducted stealth attacks, reconnaissance, intelligence-gathering and accompanied VIPs - activities that would be expected from special forces units.
Later, in January 2022 and a month before Putin ordered the full-scale invasion of Ukraine, The Times reported that 400 Wagner mercenaries were flown from the Central African Republic tasked with assassinating Ukraine's President Zelensky.
The plan was for the mercenaries to take control of the government in Kyiv and lay the groundwork for Russia to install a puppet government in its place.
By March 3, The Times reported that the Ukrainian leader had survived three assassination attempts - two of which were orchestrated by Wagner.
The group was also accused of being responsible for war crimes in the region surrounding Kyiv, with reports of torture and executions of prisoners of war.
It was also implicated in the Bucha massacre where 419 civilians were killed.
However, as with Russia's other supposed 'elite' units, Wagner's role over the course of the war has shifted from attempting clandestine missions to being used as a blunt instrument to try and overwhelm Ukrainian defences.
After pulling back from Kyiv, Russia refocused its efforts on capturing territory in the east of Ukraine.
Moscow's armies made steady advances, but were brought to a halt around the city of Bakhmut, where fighting continues to this day.
In late 2022, Wagner's leader Yevgeny Prigozhin claimed his troops - and his troops alone - were responsible for the capture of smaller towns around the city.
Around the same time, it began to emerge that Wagner had been recruiting convicts from Russian prisons to bolster its numbers.
With the promise of a pardon if a recruited convict served and survived for six months, as many as 50,000 signed up.
What was - a matter of months ago - seen as an elite, off the books unit capable of carrying out missions Russia's army otherwise could not, Wagner's forces instead became cannon fodder for Moscow's military.
Wagner boss Yevgeny Prigozhin poses with Wagner fighters in Soledar, in a video in which he claimed his forces had seized the small town near Bakhmut
Fighting for the city descended into bloodshed, with Ukrainians likening the battle for Bakhmut to a First World War 'meatgrinder' as Wagner crashed thousands of its convict troops against a wall of machine guns and artillery strikes.
After months of brutal fighting, they had been trying in vain to capture since last summer.
Prigozhin said they would withdraw on May 10 - ending their involvement in the longest and bloodiest battle of the war - because of heavy losses and inadequate ammunition supplies.
He asked defence chiefs to insert regular army troops in their place.
'I declare on behalf of the Wagner fighters, on behalf of the Wagner command, that on May 10, 2023, we are obliged to transfer positions in the settlement of Bakhmut to units of the defence ministry and withdraw the remains of Wagner to logistics camps to lick our wounds,' Prigozhin said in a statement.
'I'm pulling Wagner units out of Bakhmut because in the absence of ammunition they're doomed to perish senselessly.'
Earlier on Friday he appeared in a video surrounded by dozens of corpses he said were Wagner fighters, and was shown yelling and swearing at Defence Minister Sergei Shoigu and Chief of General Staff Valery Gerasimov.
'We have a 70% shortage of ammunition. Shoigu!
Gerasimov! Where is the ******* ammunition?' he shouted into the camera. His tirade contained a torrent of expletives that were bleeped out by his press service.
What next for special forces in Ukraine?
Prigozhin's announcement comes at a key juncture in the war, with Ukraine expected to launch a long-anticipated counter-offensive imminently.
There have already been signs that Ukrainian special forces are once again spearheading the attack, with reports last month saying the highly-trained units had already crossed the Dnieper river, probing for weak points in Russian fortifications.
Should Ukraine use the same tactics as it did in its dramatic counteroffensives last year, then we can expect to hear more whispers of Ukrainian commando units carrying out dramatic attacks behind enemy lines.
Russia, meanwhile, is facing the prospect of a military force that is increasingly made up of poorly trained reservists.
If it continues on the path of placing its most elite units on the frontlines, then even more are expected to die.